Replacing R1M2 at higher elevations (not without some transitional communities), these short, grazed swards occupy deep, acid, humid but aerated soils, often in a mosaic, generated by differences in grazing pressure and snow pack duration, with S23y, S33z and S421 and, at lower elevations and under more submediterranean, summer-dry climates, with S412. They can also occur in limestone massifs where deep, intensely leached soils develop, like in moraines, colluvia or dolines filled by the clays released by karstification. As explained under R43y, we create a new level IV unit since not all our Nardus grasslands fit confortably in R435 as R43521 (“Cantabrio-Cordilleran oro-Mediterranean mat-grass swards).